They were the most successfull of several offshoot branches of House Stark who arose in the area over the centuries. They reigned for 500 years until they made the mistake of joining House Bolton's rebellion against the King in the North, which was crushed by Karlon Stark.
It was constructed by King Jon Stark to defend the mouth of the White Knife against sea raiders (possible Iron Islanders). it is older than White Harbor. After the fall of the Greystarks, the Wolf's Den was held by many different houses.
- A Dance with Dragons 1: Dreans and Dust pages 225, 284, 448.